Many years ago, Megatronus was a brave and valiant gladiator on Cybertron, while his brother, Orion Pax, was a clerk in the Iacon Hall of Records. When the Decptiocn Galvatron appeared and the Great War began, Orion and Megatronus both entered the Autobot Army as soldiers and were trained by the great Autobot general, Sentinel Prime. One day, during a routine space mission to repair a damaged space bridge, Orion and Megatronus were seemingly lost when the bridge was attacked by a Decepticon team of soldiers. Although Orion was found and reformatted into Optimus Prime, Megatronus seemed lost forever, and the Autobots would forever mourn his death.