The older species, T. antiquus, T. natans and T. littoralis, show indications of partial adaptation to grazing, with little evidence of transverse mandibular movement while eating, and abundant dental striae indicating ingestion of sand from feeding on vegetation stranded on beaches or in shallow (less than 1 m deep) water. The later species, T. carolomartini and T. yaucensis were apparently specialized grazers that fed in deeper water; they display distinct evidence of transverse mandibular movement and lack dental striae.