By the Second Tiberium War, GDI's APCs had been completely redesigned. This new generation was a wheeled vehicle, with its machine gun removed to increase speed, especially over water. Although unable to fire at enemy ground units, the APC still retained its speed, and could travel through water, allowing GDI to quickly move squads of infantry over bodies of water and assault island fortresses, as was the case with Vega. Right after the Firestorm Crisis, the Amphibious APC was replaced by the Guardian APC and its Mobile Repair Transport variant.