Optimus Prime stands, depressed, looking out onto Cybertron. As Prowl, watching him, reflects, Prime had lost his ability to transform shortly after the Last Autobot had reconstructed him from Hi-Q's body, a result of HiQ's being "soaked" with Nucleon when he accelerated Grimlock's reconstruction. Of the "Nucleon generation" - those revived by Grimlock with Nucleon, plus Prime and Grimlock himself - Prowl was one of the few still able to transform, while the ashamed Grimlock had gone into hiding and most others, like Prime, were depressed and felt emasculated by the handicap.