Jess Squirrel was the wife of Mr. Squirrel and the mother of Sam Squirrel. She and her family lived in Mossflower when Redwall Abbey came under attack by Cluny the Scourge. Jess and her family came to Redwall after Sam led Matthias there, and they decided to remain there while the battle for Redwall waged. During another invasion from Cluny's army, Jess dropped a large barrel filled with angry bees, releasing them upon the rats.