Uganu was the Protector of Ice at the time of the Battle of the Mask Makers, serving alongside Mamuk, Agarak, Owaki, Kerato, and Etoku. A staunch ally of Ekimu, Uganu received a new Protector Mask from him at the Festival of Masks, after which the Protectors and Ekimu learned of Makuta's efforts to create a mask that combined multiple Elements. After seeing the Elemental Creatures, Ekimu and the Protectors traveled to the Temple of Time, where Ekimu donned the Mask of Time and learned of a coming cataclysm and the eventual arrival of the Toa on Okoto. With the exception of Agarak, all the Protectors then set out on a journey to obtain Elemental Crystals from the Creatures in order to power the Golden Masks that would later be worn by the Toa, with Uganu receiving a crystal from Melum.