Because the Archie Comics-published comic series Teenage Mutant Ninja Turtles Adventures originated as more or less a straight adaptation of the 1987 animated series, Splinter once again originated as Hamato Yoshi, shidoshi of the Foot Clan. As in the animated series, Hamato Yoshi loved Renaissance art, was tricked into being kicked out of the Foot Clan by Oroku Saki, and eventually mutated into a rat and witnessed four turtles mutate into humanoid forms after the five of them were exposed to Mutagen.