The Hook Beetle can be upgraded twice. It can upgrade into the Quick Beetle by using two Ancient Flowers, two Hornet Larvae, one Golden Skull, and 50 Rupees at the Scrap Shop in the Skyloft Bazaar. The Quick Beetle can then be upgraded into the Tough Beetle by using three Ancient Flowers, four Amber Relics, one Blue Bird Feather, one Goddess Plume, and 50 Rupees.