The film begins with Jak and Daxter, the protagonists, and Samos, their guardian, watching as Keira, Samos’ daughter and Jak’s romantic interest, study the Rift Rider and Precursor Ring that they found in the previous film. When activated, the Ring opens a portal that the four of them are flung through. They arrive in a dystopia known as Haven City, which is under the control of the ruthless Baron Praxis, locked in a war with a race of beasts called Metal Heads. Jak and Daxter are separated on arrival, and Jak is captured by Praxis’ Krimzon Guard. While Jak is imprisoned, he is subjected to experiments with Dark Eco; when exposed to Dark Eco, he turns into a mindless beast.