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An important Early Devonian (about 390 million years ago) fossil example from the of Gilboa, New York, was originally described as a member of the extinct order Trigonotarbida and named Gelasinotarbus? fimbriunguis.[2] It was later assigned to a new genus Attercopus[3] and reinterpreted as the oldest, and most primitive, example of a true spider (Araneae). This hypothesis was based on the supposed presence of unique spider features such as silk-producing spinnerets and the opening of a venom gland on the fang of the chelicera.

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