It falls upon April, a Shifter who can move between the worlds, to restore the Guardian and save the Balance (helped by her sidekick Crow). However, powerful forces contrive to stop her, with the Chaos Vortex, the Vanguard and their leader Jacob McAllen first among them. On her side are Cortez and the White Dragon. The entire story is narrated by Lady Alvane, a mysterious figure whose tale brackets the game. April's journey takes her through a multitude of locations across the twin worlds and beyond, leading up to a climactic but ambiguous ending, which made fans eager for a sequel.